pour clem
il existe un grand flou autour de grandis et de nombreux changement ont lieu régulièrement ci dessous la dernière version du travail des botanistes
aujourd'hui subsp evansiana n'existe plus c'est un syn. de grandis subsp grandis
Begonia grandis Dryander, Trans. Linn. Soc. London 1:
163. 1791.
秋海棠 qiu hai tang
Herbs, deciduous. Tubers subglobose, 8–12 mm in diam.
Stems 28–60 cm tall, subglabrous. Leaves cauline; stipules caducous,
oblong to lanceolate, ca. 10 × 2–4 mm, membranous,
margin entire, apex acuminate; petiole 4–20 cm, subglabrous;
blade broadly ovate, asymmetric, 10–20 × 7–14 cm, venation
palmate, 7(–9)-veined, base obliquely cordate, margin irregularly
serrulate, apex acuminate to long acuminate. Inflorescences
terminal and axillary, 7–16 cm tall, glabrous, terminal inflorescence
racemose at base, axillary inflorescence cymose; bracts
caducous, oblate to oblong, 5–13 × 2–17 mm, margin entire,
apex obtuse. Staminate flowers: pedicel 0.8–2.2 cm; tepals 4,
white to pink, glabrous, outer 2 broadly ovate or suborbicular,
1.1–1.7 × 1–1.7 cm, inner 2 obovate to obovate-oblong, 7–11 ×
3–9 mm; stamens 28–140; filaments to 6.1 mm; anthers obovoid,
0.8–1.1 mm, apex emarginate. Pistillate flowers: pedicel
2.5–5 cm; tepals 3, white to pink, glabrous; outer 2 suborbicular
or oblate-orbicular, ca. 1.2 × 1.2 cm, inner 1 obovate, ca. 8 ×
6 mm; ovary glabrous, 3-loculed; placentae axile, bilamellate;
styles fused at base or free; stigmas 2-cleft or U-shaped,
spiraled or not. Capsule pendulous, oblong, 10–12 × ca. 7 mm,
unequally 3-winged; abaxial wing oblong or triangularly oblong,
ca. 1.8 cm; lateral wings narrowly triangular, or all wings
narrowly triangular. Fl. Jul, fr. Aug. 2n = 26*.
● Forests, evergreen broad-leaved forests on montane slopes,
streamsides in dense valley forests, moist rocky cliffs of valleys, wet
fissures of limestone rocks, on rocks in shaded moist environments, on
slopes, in valleys, or by streams; 100–3400 m. Anhui, Fujian, S Gansu,
Guangxi, Guizhou, Hebei, Henan, Hubei, Hunan, Jiangxi, Shaanxi,
Shandong, Shanxi, Sichuan, Yunnan, Zhejiang.
1a. Styles free, not branched; stigma apical
or reniform; filaments connate not
beyond 1 mm ..................................... 58c. subsp. holostyla
1b. Styles connate at base or slightly connate,
branched; stigma spirally tortuous,
U-shaped; stamens more numerous,
filaments connate to or beyond 2 mm
above base.
2a. Stem weak; leaf blade abaxially
pale, occasionally reddish,
elliptic-ovate to triangular-ovate,
5–12(–20) × 3.5–9(–13) cm;
filaments connate for less than
2 mm ........................................... 58b. subsp. sinensis
2b. Stem stout; leaf blade abaxially red
or at least red on veins, ovate to broadly
so, larger than above; filaments
connate beyond 2 mm ................. 58a. subsp. grandis
58a. Begonia grandis subsp. grandis
秋海棠(原亚种) qiu hai tang (yuan ya zhong)
Begonia discolor R. Brown; B. erubescens H. Léveillé; B.
evansiana C. Andrews; B. grandis subsp. evansiana (C. Andrews)
Irmscher.
Stem stout. Leaf blade abaxially red or at least red on veins,
ovate to broadly so, larger. Filaments connate beyond 2 mm.
Capsule with evident, narrow, triangular wings.
● Streamsides in dense valley forests, moist rocky cliffs of valleys,
on rocks by streams; 100–1100 m. Anhui, Fujian, Guangxi, Guizhou,
Hebei, Henan, Hunan, Jiangxi, Shaanxi, Shandong, Shanxi, Sichuan,
Zhejiang.
Begonia grandis var. unialata Irmscher (Mitt. Inst. Allg. Bot.
Hamburg 10: 493. 1939) differs in having a capsule with 1 large wing,
and the other 2 wings very narrow or nearly absent.
58b. Begonia grandis subsp. sinensis (A. Candolle) Irmscher,
Mitt. Inst. Allg. Bot. Hamburg 10: 494. 1939.
中华秋海棠 zhong hua qiu hai tang
Begonia sinensis A. Candolle, Ann. Sci. Nat., Bot., sér. 4,
11: 125. 1859; B. bulbosa H. Léveillé; B. martini H. Léveillé.
Stem 20–40 cm tall, weak, usually not branched. Leaf blade
abaxially pale, occasionally reddish, elliptic-ovate or triangularovate,
5–12(–20) × 3.5–9(–13) cm, both surfaces glabrous or
subglabrous. Filaments connate for less than 2 mm. Styles connate
or slightly connate at base, branched; stigma spirally tortuous,
rarely U-shaped.
● Forests, on limestone rocks in shaded moist environments, on
slopes, or in valleys; 300–3400 m. Fujian, S Gansu, Guangxi, Guizhou,
Hebei, Henan, Jiangxi, Shaanxi, Shandong, Shanxi, E Sichuan, Yunnan,
Zhejiang.
Begonia grandis var. puberula Irmscher (Mitt. Inst. Allg. Bot.
Hamburg 10: 496. 1939) differs in having a leaf blade adaxially densely
hispidulous (hairs ca. 0.3 mm) and sparsely hirsute (hairs 0.8–0.9 mm),
while var. villosa T. C. Ku (Fl. Reipubl. Popularis Sin. 52(1): 166. 1999)
differs in having a leaf blade adaxially slightly floccose.
58c. Begonia grandis subsp. holostyla Irmscher, Mitt. Inst. Allg.
Bot. Hamburg 10: 498. 1939.
全柱秋海棠 quan zhu qiu hai tang
Stem (12–)20–40(–60) cm tall, weak, usually not branched.
Leaf blade triangular-ovate, 3–13 × 2–10 cm. Styles free,
not branched; stigmas capitate or reniform. Fl. Jul–Aug, fr. Sep–
Oct.
● Evergreen broad-leaved forests on montane slopes, on shaded
moist rocks of shrubberies, wet fissures of limestone rocks, moist places
by rocks; 2200–2800 m. Sichuan (Muli), Yunnan.